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I think VE = motors ability to pump a volume of air / theoretical volume pumped. If you modify the internals of said motor(pump) you are changing that value. For example, a lower flow rate due to restriction in intercooler ends up reducing your motors ability to pump a volume of air (VE) and bigger exhaust may increase your motors ability to pump a volume of air.
I have heard alot of people (not here) say things like a turbo or charged motor is more than 100% VE, and I have to disagree, ignoring losses in things like cam overlap etc. All we are doing is increasing the density of a given volume of air...
VE is a function of many less variable (mechanical) things like cams, port size etc. Once an engine is defined mechanically, VE changes mostly with RPM, however i think, the flatter the torque curve, the less the change is. For example, my Varioram motor attempts to adjust intake runner sizes to better maximize the VE of the Engine at specific RPMs.
In this application, I see 2 pumps in sequence. If the first pump (SC) is more efficient the pressure between them goes up, if the second is more efficient the pressure drops.
If I had to stab at his situation here I think either his motor is really waking up at mid RPM and up and really beating the pants off the SC. Most would say this is due to some mechanical items, cams, port size etc. I see an opportunity, if we were to increase the output of the SC (bigger) we could see bigger mass air flow at high RPM...
I certainly don't want anyone to think I am flaming, in fact I am amazed at the amount of help i see in this thread, I am hoping to learn...
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